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Introductory

WILDLIFE SPECIES: Martes americana | American Marten
ABBREVIATION : MAAM COMMON NAMES : American marten marten pine marten Pacific marten eastern marten western marten Rocky Mountain marten TAXONOMY : The currently accepted scientific name for the American marten is Martes americana. Five subspecies are recognized: Martes americana ssp. americana, eastern marten M. americana ssp. actuosa, marten M. americana ssp. atrata, Newfoundland pine marten M. americana ssp. origenes, Rocky Mountain marten M. americana ssp. sierrae, western marten The name Martes caurina has been used in earlier literature [7,21]. ORDER : Carnivora CLASS : Mammal FEDERAL LEGAL STATUS : No special status OTHER STATUS : The American marten has been eliminated from much of its former range in the eastern United States and Canada. It is limited to small, isolated areas of the Southwest [7,12,21]. The Newfoundland pine marten is listed as threatened in Newfoundland [22]. COMPILED BY AND DATE : S. A. Snyder, April 1991 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Snyder, S. A. 1991. Martes americana. In: Remainder of Citation

WILDLIFE DISTRIBUTION AND OCCURRENCE

WILDLIFE SPECIES: Martes americana | American Marten
GENERAL DISTRIBUTION : American Martens occur throughout much of the northern boreal forest of Canada and in isolated portions of northeastern United States and Canada. They also occur in isolated areas of the Pacific Coast States, the inland Rocky Mountain West, and the Southwest. American martens are rare in the eastern United States but can be found in limited areas of Wisconsin, Michigan, and Minnesota [1,3].

American martens are also found on the following Alaskan islands: Admiralty, Kuiu, Kupreanof, and Revillagigedo. They have been stocked on Prince of Wales, Garanof, and Chichagof islands. Subspecies: Eastern marten inhabit eastern North America. Western marten inhabit portions of California and Nevada. Rocky Mountain marten inhabit the western United States [7,21]. ECOSYSTEMS : FRES11 Spruce-fir FRES18 Maple-beech-birch FRES19 Aspen-birch FRES23 Fir-spruce FRES24 Hemlock-Sitka spruce STATES :
AK AR CA CO ID MI MN MT NY OH UT WA WY

AB BC MB NB NF NT NS ON PE PQ
SK YK
BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 4 Sierra Mountains 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau KUCHLER PLANT ASSOCIATIONS : K001 Spruce - cedar - hemlock forest K002 Cedar - hemlock - Douglas-fir forest K003 Silver fir - Douglas-fir forest K004 Fir - hemlock forest K005 Mixed conifer forest K007 Red fir forest K008 Lodgepole pine - subalpine forest K013 Cedar - hemlock - pine forest K014 Grand fir - Douglas-fir forest K015 Western spruce - fir forest K020 Spruce - fir - Douglas-fir forest K021 Southwestern spruce -fir forest K093 Great Lakes spruce - fir forest K094 Conifer bog K096 Northeastern spruce - fir forest K102 Beech - maple forest K107 Northern hardwoods - fir forest K108 Northern hardwoods - spruce forest SAF COVER TYPES : 5 Balsam fir 12 Black spruce 13 Black spruce - tamarack 22 White pine - hemlock 25 Sugar maple - beech - yellow birch 30 Red spruce - yellow birch 31 Red spruce - sugar maple - beech 32 Red spruce 33 Red spruce - balsam fir 34 Red spruce - Fraser fir 201 White spruce 204 Black spruce 205 Mountain hemlock 206 Engelmann spruce - subalpine fir 207 Red fir 211 White fir 212 Western larch 213 Grand fir 215 Western white pine 216 Blue spruce 218 Lodgepole pine 223 Sitka spruce 224 Western hemlock 225 Western hemlock - Sitka spruce 226 Coastal true fir - hemlock 227 Western redcedar - western hemlock 228 Western redcedar 229 Pacific Douglas-fir 230 Douglas-fir - western hemlock 232 Redwood 243 Sierra Nevada mixed conifer 251 White spruce - aspen 253 Black spruce - white spruce SRM (RANGELAND) COVER TYPES : NO-ENTRY PLANT COMMUNITIES : American martens are found in climax coniferous and mixed forests. They need dense overstory (>30 percent) and sufficient understory cover for hiding and denning. Mesic sites that support dense, succulent understory vegetation for American marten prey species are considered the best habitat [12]. During the summer, American martens may hunt for food in open meadows bordering dense forests if hiding cover is present [1,11,12]. American martens inhabit high elevation basins in spruce (Picea spp.)-subalpine fir (Abies lasiocarpa) or mountain hemlock (Tsuga mertensiana) forests in the West. Mature lodgepole pine (Pinus contorta) stands that include spruce or subalpine fir will also support American marten. Although American martens are usually found at high elevations, they will use forests at lower elevations with high precipation, such as cedar (Thuja spp.)-grand fir (Abies grandis) [12]. In the East they are found in maple (Acer spp.)-beech (Fagus spp.)-birch (Betula spp.) forests. REFERENCES : NO-ENTRY

BIOLOGICAL DATA AND HABITAT REQUIREMENTS

WILDLIFE SPECIES: Martes americana | American Marten
TIMING OF MAJOR LIFE HISTORY EVENTS : Minimum breeding age - 1 year for males 2 years for females Breeding season - late July to August Gestation period - active gestation is 27 days following delayed implantation Birthing season - April, two to four young Maximum lifespan - 17 years in captivity PREFERRED HABITAT : American martens prefer dense, old-growth conifer and mixed stands. Stands must have sufficient understory to support various rodents, such as mice (Cricetids) and voles ( Microtines), the major food source. Martens usually den in rotten logs. They may also den in rock slides and slash piles [4]. COVER REQUIREMENTS : Cover requirements for American martens include dense conifer or mixed forests with 40 to 60 percent canopy closure. They will avoid areas that are so dense that herbaceous cover is suppressed [12]. Dense understory, including slash or rotten logs and stumps, is necessary for denning and hiding [18]. Open areas adjacent to these forests will be used for hunting only if they provide adequate hiding cover and food. Uneven-aged stands are most beneficial because their vegetation is more diverse which leads to a greater food base [1,11,12]. FOOD HABITS : American martens prey on small mammals, particularly mice and voles. Other small mammal prey include ground squirrels (Spermophilus spp.), flying squirrels (Glaucomys spp.), chipmunks (Eutamias spp.), and snowshoe hares (Lepus americanus). The American marten's diet can also include insects, various fruits and nuts, and passerine birds [1,14]. PREDATORS : Amer5ican marten predators include man, bear (Ursus spp.), mountain lion (Felis concolor), lynx (F. lynx), bobcat (Lynx rufus), coyote (Canis latrans), gray wolf (C. lupus), eagles (Accipitrines), and the great-horned owl (Bubo virginianus). MANAGEMENT CONSIDERATIONS : Koehler and others [12] listed some management recommendations for the American marten. During winter American marten need old-growth forests on mesic sites with a canopy cover exceeding 30 percent. In winter, they rarely cross openings greater than 300 feet (90 m) and do not hunt in openings. Mature forests may support more American marten in certain areas, but diverse forest communities support more American marten over time. In summary, openings should be small, undisturbed, old-growth forests on mesic sites, and forest diversity must be maintained. American martens can tolerate human activity; however, road closures after logging are important to ensure the geographical isolation that American martens require. REFERENCES : NO-ENTRY

FIRE EFFECTS AND USE

WILDLIFE SPECIES: Martes americana | American Marten
DIRECT FIRE EFFECTS ON ANIMALS : American martens could be trapped in a rapidly spreading, intense fire. However, direct fire mortality probably has little effect on the population as a whole. HABITAT RELATED FIRE EFFECTS : Fires that completely consume the understory and/or reduce the canopy closure to less than 30 percent are detrimental to American marten populations in the short-term. However, fires that create a mosaic of diverse habitats provide the best cover for American marten and their prey in the long-term. Marten populations tend to increase several decades after fire as adequate food and cover are replaced [11,12,20]. Koehler and others [12] suggested that low-intensity fires on mesic sites where canopy cover is maintained at greater than 30 percent may not adversely affect American marten habitat. FIRE USE : Prescribed fire can be used to create a diversity of forest communities, which, over time, may support more American marten. Fire can create and maintain openings where abundant fruits, insects, ground squirrels, and voles provide summer food. Smaller burns may be more beneficial because they are less restrictive to American marten movement in winter, and cover is reestablished more quickly. Islands of climax forest containing logs, stumps, and slash for denning should always be maintained. Islands of forest should be no farther apart than 55 yards (50 m) [11,12]. REFERENCES : NO-ENTRY

References for species: Martes americana


1. Allen, Arthur W. 1984. Habitat suitability index models: marten. FWS/OBS-82/10.11 (Revised). Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 13 p. [11714]
2. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434]
3. Burt, William H.; Grossenheider, Richard P. 1976. A field guide to the mammals. Boston, MA: Houghton Mifflin. 289 p. [13685]
4. Buskirk, Steven W.; Forrest, Steven C.; Raphael, Martin G.; Harlow, Henry J. 1989. Winter resting site ecology of marten in the central Rocky Mountains. Journal of Wildlife Management. 53(1): 191-196. [6896]
5. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905]
6. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998]
7. Hall, E. Raymond. 1946. Mammals of Nevada. Berkeley, CA: University of California Press. 710 p. [13688]
8. Heinselman, Miron L. 1973. Fire in the virgin forests of the Boundary Waters Canoe Area, Minnesota. Quaternary Research. 3: 329-382. [282]
9. Honacki, James H.; Kinman, Kenneth E.; Koeppl, James W., eds. 1982. Mammal species of the world. Lawrence, KA: Allen Press Inc. 694 p. [13703]
10. Ingram, Rod. 1973. Wolverine, fisher, and marten in central Oregon. Central Oregon Adminstrative Report No. 73-2. Salem, OR: Oregon State Game Commission. 41 p. [13472]
11. Koehler, Gary M.; Hornocker, Maurice G. 1977. Fire effects on marten habitat in the Selway-Bitterrroot Wilderness. Journal of Wildlife Management. 41(2): 500-505. [7637]
12. Koehler, Gary M.; Moore, William R.; Taylor, Alan R. 1975. Preserving the pine martin: management guidelines for western forests. Western Wildlands Summer 1975. Missoula, MT: University of Montana, Montana Forest and Conservation Experiment Station. 6 p. [8452]
13. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384]
14. Lensink, Calvin J.; Skoog, Ronald O.; Buckly, John L. 1955. Food habits of marten in interior Alaska and their significance. Journal of Wildlife Management. 19(3): 364-368. [13689]
15. Mech, L. David; Rogers, Lynn L. 1977. Status, distribution, and movements of martens in northeastern Minnesota. Res. Pap. NC-143. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Forest Experiment Station. 7 p. [13465]
16. Meehan, William R. 1974. The forest ecosystem of southeast Alaska: 4. Wildlife habitats. Gen. Tech. Rep. PNW-16. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 32 p. [13479]
17. Rowe, J. S.; Scotter, G. W. 1973. Fire in the boreal forest. Quaternary Research. 3: 444-464. [72]
18. Spencer, Wayne D. 1987. Seasonal rest-site preferences of pine martens in the northern Sierra Nevada. Journal of Wildlife Management. 51(3): 616-621. [13690]
19. Thompson, Ian D. 1991. Could marten become the spotted owl of eastern Canada? Forestry Chronicle. 67(2): 136-140. [7553]
20. Viereck, Leslie A.; Schandelmeier, Linda A. 1980. Effects of fire in Alaska and adjacent Canada--a literature review. BLM-Alaska Tech. Rep. 6. Anchorage, AK: U.S. Department of the Interior, Bureau of Land Mangement, Alaska State Office. 124 p. [7075]
21. Warren, Edward Royal. 1942. The mammals of Colorado. Norman, OK: University of Oklahoma Press. 330 p. [13687]
22. Committee on the Status of Endangered Wildlife in Canada. 1992. Canadian species at risk. Ottawa, ON. 10 p. [26183]


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Information Courtesy: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory. Fire Effects Information System

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